Nicholas Gladman


I joined the University of Leeds in 2014 as a PhD student investigating the energetics and mechanics of European common cuttlefish (Sepia officinalis) jet propulsion swimming (Supervised by Dr Graham Askew). Prior to this I undertook a BSc in Aquatic Zoology at the University of Hull (2009-2012) and an MRes in Animal Behaviour at the University of Nottingham (2012-2014). 


My undergraduate work (supervised by Dr Lesley Morrell) focused on the energetic costs of facultative colour change in Trinidadian guppies (Poecilia reticulata), revealing frequent changes in colouration appeared to carry an energetic cost as measured through increased food consumption rates. During my MRes I moved to investigate the stridulatory apparatus of corixids (water boatmen; Supervised by Dr Tom Reader), this research found populations where sister species co-habit showed structural differences in the stridulatory apparatus, with this likely resulting from plasticity in growth of these structures, perhaps as a result of competition driving animals to alter their diet with resultant impacts upon the calls of males.  

Research interests

My current research looks at the energetics of European cuttlefish (Sepia officinalis) jet propulsion swimming. Looking at changes from the biochemical level through to whole animal swimming mechanics, enabling the efficiency of the whole propulsive system to be investigated. 


My research interests lie predominantly in the biomechanics and ecophysiology of aquatic organisms, I am particularly interested in understanding how an animal’s physiology, environment and behaviour interact. 



  • BSc (Hons) Aquatic Zoology
  • MRes Animal Behaviour

Research groups and institutes

  • Ecology and Evolution
  • Sport and Exercise Sciences