John Grahame


BSc (Zoology), University of the West Indies (Mona).

PhD, Marine Science Laboratories, University College of North Wales.

Now retired from Senior Lectureship, School of Biology, University of Leeds.

Research interests

Biogeography of the 'rough periwinkles' (Littorina species); measurement, quantification of variation and its interpretation among turbinate snails using Littorina as an example.

<h4>Research projects</h4> <p>Any research projects I'm currently working on will be listed below. Our list of all <a href="">research projects</a> allows you to view and search the full list of projects in the faculty.</p>

Professional memberships

  • The Malacological Society of London (current president).