Postdoctoral Research Assistant (1978 – 1984), Universities of Cambridge & Oxford
Independent Research Fellow (1984 – 1994), University College Cardiff & University of Leeds.
Lecturer (1994 – 1997), University of Leeds
Reader in Integrative Neuroscience (1997 - 2009), University of Leeds.
Visiting Senior Research Fellow (2009 - ), University of Leeds
Research interests
The central cholinergic system of the mammalian brain
Characterisation of some unusual members of the Rhizaria (Foraminifera, Cercozoa, Gromia) and the Alveolata (dinoflagellata) that inhabit the coastal areas, freshwater lochs and bogs of north west Scotland
<h4>Research projects</h4>
<p>Any research projects I'm currently working on will be listed below. Our list of all <a href="https://biologicalsciences.leeds.ac.uk/dir/research-projects">research projects</a> allows you to view and search the full list of projects in the faculty.</p>
Qualifications
1973: BSc. (Honours, Class II/I) Physiology, University College London.
1978: PhD in Neuroscience, Churchill College, Cambridge University.
2017: BSc. (Honours, Open, Class II/I) on topics in theoretical physics, Open University.
Professional memberships
The Society for Neuroscience (Emeritus member)
The Society of Authors
The Scottish Association for Marine Science (Clan member)