Professor Susan A. Ward


Professor Ward gained her DPhil in Physiology from Oxford University in 1974. Following two years as a Lecturer in the Department of Physiology at Liverpool University, she moved to the University of California at Los Angeles, becoming Professor of Anesthesiology and Physiology in 1988. She returned to the United Kingdom in 1993, first to the Department of Physiology at the University of London's St George's Hospital Medical School, then to South Bank University as Chair of Sports Science and, in 1998, to the University of Glasgow as Director of the Centre for Exercise Science and Medicine. In 2003, she was appointed to the Chair of Sport and Exercise Science and Head of the School of Sport and Exercise Sciences at the University of Leeds, and then Emeritus Professor in 2007. With some 180 publications, her research interests include the control of ventilation, pulmonary gas exchange and muscle energetics during exercise in health, altered environments and disease; and the methodology and applications of cardiopulmonary exercise testing. She is a Fellow of the American College of Sports Medicine and the European Respiratory Society. She is immediate past Editor-in-Chief of the "European Journal of Applied Physiology"; previously served as Chair of the Pulmonary Circulation, Gas Exchange and Exercise Group of the European Respiratory Society; and is a consultant for Xtreme Everest 2 (, a collaborative venture between Duke University USA, the University of Southampton and University College London.

Research interests

Recent publications:

Berger, K., S. Fagondes, R. Giugliani, K. Hardy, C. McArdle, D. Rapoport, M. Scarpa, M. Tobin and S.A. Ward. Respiratory and sleep disorders in mucopolysaccharidosis. J. Inherit. Metab. Dis. 36:201-210, 2013.

Cannon, D.T., F.A. Howe, B.J. Whipp, S.A. Ward, D.J. McIntyre, C. Ladroue, J.R. Griffiths, G.J. Kemp and H.B. Rossiter. Muscle metabolism and activation heterogeneity by combined 31P chemical shift and T2 imaging, and pulmonary O2 uptake during incremental knee-extensor exercise. J. Appl. Physiol. 115: 839-849, 2013.

Ward, S.A. Invited Commentary on "Mechanism of augmented exercise hyperpnea in chronic heart failure and dead space loading" by Poon and Tin. Respir Physiol Neurobiol. 189:  203-210, 2013 [F1000Prime recommended]. 

Whipp, B.J., and S.A. Ward. The respiratory system. In: History of Exercise Physiology. Ed: C. Tipton. Human Kinetics: Champaign, pp. 211-245, 2014.

Ward, S.A. Regulation of breathing during exercise. In: Colloquium Series on Integrated Systems Physiology: The Respiratory System. Eds: Granger, D.N. and J.P. Granger. Morgan & Claypool Life Sciences Publishers: Princeton, 2014.

Murgatroyd, S.R., Wylde, L.A., Cannon, D.T., Ward, S.A., and H.B. Rossiter. A ‘ramp-sprint’ test characterises the indices of aerobic function and exercise intensity domains in humans in a single laboratory visit. Europ. J. Appl. Physiol. 114: 1863-1874, 2014.

Ward, S.A. Commentary on Viewpoint: The two-hour marathon: what's the equivalent for women? J. Appl. Physiol. 118: 1328, 2015.

Ward, S.A. Commentary on Viewpoint: Precedence and autocracy in breathing control. J. Appl. Physiol. 118(12): 1559, 2015.

Ward, S.A. Back to Basics: The Fundamental Principles of Exercise Physiology. Acta Physiol 217 (Suppl. 708): 11-12, 2016.

Puente-Maestu, L., P. Palange, R. Casaburi, P. Laveneziana, F. Maltais, J.A. Neder, D. O’Donnell, P. Onorati, J. Porszasz, R. Rabinovich, H.B. Rossiter, S. Singh, T. Troosters and S.A. Ward. Use of exercise testing in the evaluation of interventional efficacy: an official ERS statement. Eur. Resp. J. 47: 429-460, 2016.

Laveneziana, P., A. Neder, S.A. Ward, and V. Poberezhets. Clinical exercise testing: basic principles and practice. Breathe 13(3):163-164, 2017.

Ward, S.A., M.P.W. Grocott and D.Z.H. Exercise testing, hypoxia, and supplemental oxygen. Ann Am Thorac Soc 14(suppl 1): S140-S148, 2017

Levett, D.Z.H., Jack, S., Swart, M., Carlisle, J., Wilson, J., Snowden, C., Riley, M., Danjoux, G., Ward, S.A., Older, P., Grocott, M.P.W. and the UK Perioperative Exercise Testing and Training Society. Perioperative Cardiopulmonary Exercise Testing for Non-cardiopulmonary Surgery: Consensus Clinical Guidelines. Br. J. Anaesth. 120: 484-500, 2018.

Ward, S.A. Open-circuit respirometry: real-time, laboratory based systems. Europ. J. Appl. Physiol. 118: 875-898, 2018.

Palange, P, Laveneziana, P, Neder, A, and Ward, S.A. Introduction: CPET in clinical practice. Recent advances, current challenges and future directions. In: Clinical Exercise Testing (ERS Monograph). Eds.: Palange P, Laveneziana P, Neder JA, Ward SA. Sheffield, European Respiratory Society, 2018; pp. x–xxv.

Ward, S.A. Determinants of the physiological systems’ responses to muscular exercise in healthy subjects. In: Clinical Exercise Testing (ERS Monograph). Eds.: Palange P, Laveneziana P, Neder JA, Ward SA. Sheffield, European Respiratory Society, 2018; pp. 1–33.

Ward, S.A. Respiratory physiology. In: ERS Handbook on Adult Respiratory Medicine, 3rd ed. Eds: Palange P., and A. Simonds. ERS: Sheffield UK, pp.28-40, 2019.

Whipp, B.J., and S.A. Ward Control of ventilation. In: ERS Handbook on Adult Respiratory Medicine, 3rd ed. Eds: Palange P., and A. Simonds. ERS: Sheffield UK, pp. 41-47, 2019.

Levett, D. S.A. Ward and M. Grocott. Exercise physiology at altitude. Current Opinion in Physiology (Exercise Physiology), 10:210–218, 2019.

Ward S.A. Control of the exercise hyperpnea. Current Opinion in Physiology (Exercise Physiology), 10:166–172, 2019.

Ward, S.A., and B.J. Whipp. Exercise physiology. In: . Ed: S. Janes. Elsevier, 2019. In press.


<h4>Research projects</h4> <p>Any research projects I'm currently working on will be listed below. Our list of all <a href="">research projects</a> allows you to view and search the full list of projects in the faculty.</p>


  • DPhil, Physiology, Oxford University

Professional memberships

  • American Physiological Society
  • American College of Sports Medicine
  • The Physiological Society
  • Perioperative Exercise Testing and Training Society